Reading and Writing Module 2 (Hard) Question 4

Some scientists have believed that pollen-carrying insects were not very _____, but according to paleoentomologist Alexander Khramov, insects were carrying pollen between plants long before the advent of mammals. Six of 425 Tillyardembia fossils studied by Khramov’s team in Moscow contained pollen grains inside the insects, a finding that pushes back the oldest fossil record of pollinators by 120 million years.
Which choice completes the text with the most logical and precise word or phrase?

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